Abstract

Taxonomic changes in the New World Hydnocerina (Coleoptera: Cleridae: Clerinae: Hydnocer¬ini) at the species and genus level are presented. Phyllobaenus suturalis (Klug, 1842) and Hydnocera limbata Spinola, 1844 are designated junior synonyms of P. pallipennis (Say, 1825), new synonymy. Phyllobaenus nigrescens (Schaeffer, 1909) is designated a junior synonym of P. verticalis (Say, 1835), new synonymy. Iso¬hydnocera curtipennis (Newman, 1840), P. lineaticollis Spinola, 1844, and Clerus brachypterus Klug, 1842, are re-established as junior synonyms of P. verticalis (Say, 1835), status reinstated, and I. curtipennis is accord¬ingly reassigned to Phyllobaenus Dejean as P. curtipennis, new combination. Cephaloclerus < i>Kuwert, 1893 is removed from synonymy under Phyllobaenus and re-elevated to genus rank, status resurrected. The genus Isohydnocera Chapin, 1917 is designated a junior synonym of Phyllobaenus Dejean, 1833, new synonymy, and I. liebecki Wolcott, 1928 is designated a junior synonym of I. tabida (LeConte, 1849), new synonymy. The new genus Neohydnocera is described to accommodate species formerly known as Isohydnocera Chapin. Hydnocera longicollis Zeigler, 1844 is removed from synonymy under I. curtipennis, status resurrected, and recognized as Neohydnocera longicollis, new combination. The following new combinations for species pre¬viously considered Isohydnocera are presented: Neohydnocera aegra (Newman, 1840), new combination, Phyllobaenus albocinctus (Horn, 1871), new combination, N. brunnea (Chapin, 1917), new combination, N. californica (Barr, 1966), new combination, N. chiricahuana (Knull, 1949), new combination, P. cryptocer¬inus (Gorham, 1883), new combination, N. gerhardi < i>(Wolcott, 1910), new combination, N. mima (Wolcott, 1928), new combination, N. nigrina (Schaeffer, 1908), new combination, N. ornata (Wolcott, 1908), new combination, N. pusilla (Schaeffer, 1909), new combination, P. schusteri (LeConte, 1866),new combina¬tion, N. tabida (LeConte, 1849), new combination, and P. tricondylae (LeConte, 1849), new combination.

References

• Barr WF. 1966. Two new species of Pacific Coast Cleridae (Coleoptera). Biological Society of Nevada Occasional Papers 10: 1–4. 
• Barr WF. 1975. Family Cleridae. Volume 4. Family 73. p. 1–18. In: Arnett RH Jr. (ed.). Checklist of the beetles of North and Central America and the West Indies. Flora and Fauna Publications; Gainesville, FL. 254 p 
• Beal JA, Massey CL. 1945. Bark beetles and ambrosia beetles (Coleoptera: Scolytoidea): with special references to species occurring in North Carolina. Duke University School of Forestry Bulletin 10(1): 1–178. 
• Bøving AG, Champlain AB. 1920. Larvae of the North American beetles of the family Cleridae. Proceedings of the United States National Museum 57: 576–649. 
• Bøving AG, Craighead FC. 1931. An illustrated synopsis of the principal larval forms of the order Coleoptera. Entomológica Americana 11(4): 275–351. 
• Chapin EA. 1917a. Miscellaneous notes on Coleoptera. Bulletin of the Brooklyn Entomological Society 12: 29–31. 
• Chapin EA. 1917b. Studies in the Hydnocerini. The hydnoceroid genera. Bulletin of the Brooklyn Entomological Society 12: 83–85. 
• Chapin EA. 1922. New North American Hydnocera (Coleoptera: Cleridae). Proceedings of the Biological Society of Washington 35: 55–58. 
• Chapin EA. 1948. Notes on Cleridae. Overgedrukt uit: Entomologische Berichten 285(12): 304. 
• Chevrolat MA. 1874. Catalogue des Clerides de la collectiòn de M.A. Chevrolat. Revue et Magasin de Zoologie (Series 3) 2(8–9): 252–329. 
• Chittenden FH. 1890. Remarks on the habits of some species of Cleridae. Entomológica Americana 6: 154-155. 
• Corporaal JB. 1950. Cleridae. In: Hinks WD (ed.). Coleopterorum catalogus supplementa, Pars 23 (editio secunda). W. Junk; The Hague, Netherlands. 373 p 
• Craighead FC. 1950. Insect enemies of eastern forests. United States Department of Agriculture, Miscellaneous Publications 657: 1–679. 
• Dejean, PFMA. 1833. Catalogue des Coléoptères de la collection de M. le Comte Dejean, livre. 2. Méquignon-Marvis; Paris. 80 p. (p. 97–176) 
• Dillon ES, Dillon LS. 1972. A manual of common beetles of eastern North America. Volume 1. Dover Publications, Inc.; New York, NY. 434 p 
• Downie NM, Arnett RH Jr. 1996. Family 73. Cleridae. p. 940–956. In: Downie NM, Arnett RH Jr. The beetles of northeastern North America. Volume II. Sandhill Crane Press; Gainesville, Florida. 1721 p 
• Drury C. 1879. List of the Coleoptera observed in the vicinity of Cincinnati. Journal of the Cincinnati Society of Natural History 2: 162–178. 
• Drury C. 1902. A revised list of Coleoptera observed near Cincinnati, Ohio. Journal of the Cincinnati Society of Natural History 20: 107–196. 
• Ekis G. 1975. Taxonomic and nomenclatural status of clerid taxa described by Massimiliano Spinola (1780–1857) (Coleoptera: Cleridae). Bollettino del Museo di Zoologia dell’ Università di Torino 1: 1–80. 
• Ekis G, Gupta AP. 1971. Digestive system of Cleridae (Coleoptera). International Journal of Insect Morphology and Embryology 1(1): 51–86. 
• Eliason EA, Potter DA. 2000. Biology of Callirhytis cornigera (Hymenoptera: Cynipidae) and the arthropod community inhabiting its galls. Environmental Entomology 29: 551–559. 
• Erichson WF. 1845. Insecta–Coleoptera. Clerii. p. 174–179. In: Reports on the progress of zoology and botany. The Ray Society; Edinburgh, UK. 348 p 
• Evans AV. 2014. Beetles of eastern North America. Princeton University Press; Princeton, NJ. 560 p 
• Felt EP. 1906. Insects affecting park and woodland trees. New York State Museum Memoirs 8(2): 333–877. 
• Gerstmeier R. 2009. Revision of the genera Diplocladus Fairmaire and Strotocera Schenkling (Coleoptera: Cleridae: Tillinae). Zootaxa 2242: 1–54. 
• Gorham HS. 1877. Descriptions of new species of Cleridae. Transactions of the Royal Entomological Society of London 18(3): 245–263. 
• Gorham HS. 1886. Insecta, Coleoptera, Cleridae. Biologia Centrali-Americana 3(2, supplement): 313–360. 
• Gosling DCL. 1980. An annotated list of the checkered beetles (Coleoptera: Cleridae) of Michigan. The Great Lakes Entomologist 13(2): 65–76. 
• Bartlett JS, Chapman EG, Cameron SL. 2013. A molecular phylogeny of the checkered beetles and a description of Epiclininae subfam. nov. (Coleoptera: Cleroidea: Cleridae). Systematic Entomology 38(3): 626–636. 
• Hamilton J. 1895. Catalogue of the Coleoptera of southwestern Pennsylvania, with notes and descriptions. Transactions of the American Entomological Society 22: 317–381. 
• Horn GH. 1871. Descriptions of new Coleoptera of the United States, with notes on known species. Transactions of the American Entomological Society 3(1870–1871): 325–344. 
• Klug JCF. 1842. Versuch einer systematischen Bestimmung und Auseinandersetzung der Gattungen und Arten der Clerii, einer Insektenfamilie aus der Ordung der Coléoptèren. Abhandlungen der Koniglichen Akademie der Wissenschaften in Berlin 1840–1842(1842): 259–397. 
• Knull JN. 1930. Notes on Coleoptera, No. 2. Entomological News 41: 82–86. 
• Knull JN. 1932. Notes on Coleoptera, No. 3. Entomological News 43: 42–45. 
• Knull JN. 1949. Three new species of Cleridae (Coleoptera). Ohio Journal of Science 49: 199–200. 
• Knull JN. 1951. The checkered beetles of Ohio (Coleoptera: Cleridae). Ohio Biological Survey Bulletin 8(42): 268–350. 
• Kolibáč J. 1998. Classification of the subfamily Hydnocerinae Spinola, 1844 (Coleoptera: Cleridae). Acta Musei Moraviae Scientiae Biologicae 83: 127–210. 
• Kuwert AF. 1893. Neue Africanische Cleriden. Annales de la Société Entomologique de Belgique 37: 486–491. 
• Leavengood JM Jr. 2008. The checkered beetles (Coleoptera: Cleridae) of Florida. M.S. Thesis, University of Florida. 206 p 
• Leavengood JM Jr. 2014. Nomenclatural and distributional notes regarding some North American species of Phyllobaenus (Coleoptera: Cleridae: Hydnocerinae). Giornale Italiano di Entomologia 13(59): 471–480. 
• Leavengood JM Jr. 2020. Phyllobaenus thomasi and P. turnbowi, two new species from Mexico and Belize (Coleoptera: Cleridae: Hydnocerinae: Hydnocerini). Insecta Mundi 0833: 1–6. 
• Leavengood JM Jr. 2024. Corrections and additions to the Hydnocerina (Coleoptera: Cleridae: Clerinae: Hydnocerini) of Mexico. Insecta Mundi 1054: 1–5. 
• Garner BH. 2014. Nomenclatural notes on some checkered beetle (Coleoptera: Cleridae) types of the British Museum of Natural History (London). Zootaxa 3760(3): 301–335. 
• Pinkerton MG, Rifkind J. 2022. Description of the new genus Tarsobaenus and three new species from Costa Rica (Coleoptera: Cleridae: Hydnocerinae). Zootaxa 5138(2): 167–176. 
• Rifkind J. 2020. Phyllobaenus inusitatotibialis, n. sp., a new species from southern Arizona (Coleoptera: Cleridae: Hydnocerinae: Hydnocerini). Proceedings of the Entomological Society of Washington 122(3): 673–677. 
• LeConte JL. 1849. Synopsis of the Coleopterous insects of the group Cleridae which inhabit the United States. Annals of the Lyceum of Natural History of New York 5(1851–52): 9–35. 
• LeConte JL. 1859. American Entomology. A description of the insects of North American by Thomas Say. (Edited by LeConte) vol. 2. Bailliere; New York, NY. 814 p. (Reprinted 1869, 1885, 1891) 
• Reprinted 1869, 1885, 1891) LeConte JL. 1866. List of the Coleoptera of North America. Smithsonian Miscellaneous Collections 6(140): 1–78. 
• Lohde R. 1900. Cleridarum Catalogus. Stettiner Entomologische Zeitung 61: 3–148. 
• Majka CG. 2006. The checkered beetles (Coleoptera: Cleridae) of the Maritime Provinces of Canada. Zootaxa 1385: 31–46. 
• Mawdsley JR. 1999. New records and biological notes on species of Cleridae (Coleoptera) from the Adirondack Park, New York. Great Lakes Entomologist 32(1–2): 39–45. 
• Mawdsley JR. 2002. Ecological notes on species of Cleridae (Coleoptera) associated with the prairie flora of central North America. Great Lakes Entomologist 35: 15–22. 
• Mawdsley JR. 2006. Taxonomy, phylogeny, and cladistic biogeography of the genus Omadius (Coleoptera: Cleridae). Contributions of the American Entomological Society 34(4): 1–98. 
• Newman E. 1838. Entomological notes. Entomological Magazine 56: 168–181. 
• Newman E. 1840. Descriptions of some new species of coleopterous insects. Annals and Magazine of Natural History (2)4: 362–368. 
• Opitz W. 2002. Family 73. Cleridae. p. 267–280. In: Arnett RH Jr., Thomas MC, Skelley PE, Frank JH (eds.). American beetles, Vol. 2. CRC Press; Boca Raton, FL. 861 p 
• Opitz W. 2010. Classification, natural history, phylogeny and subfamily composition of the Cleridae and generic content of the subfamilies. Entomologica Basiliensia et Collectionis Frey 32: 31–128. 
• Osten-Sacken CR. 1861. On the Cynipidae of North American oaks and their galls. Proceedings of the Entomological Society of Philadelphia 1: 47–72. 
• Papp CS. 1960. The Cleridae of North America, Part I: The geographical distribution of the Cleridae of North America, North of the Panama Canal. Southern California Academy of Sciences Bulletin 58(3): 78–88. 
• Peck SB, Thomas MC. 1998. A distributional checklist of the beetles (Coleoptera) of Florida. Arthropods of Florida and Neighboring Land Areas 16. Florida Department of Agriculture and Consumer Services, Gainesville. 180 p 
• Pierce WD, Cushman RA, Hood CE. 1912. The insect enemies of the cotton boll weevil. United States Department of Agriculture. Bureaus of Entomology. Technical Series 100: 1–99. 
• Sabrosky CW. 1934. Notes on the larvae and larval habits of Isohydnocera curtipennis (Newman) (Coleoptera: Cleridae). Journal of the Kansas Entomological Society 7: 65–68. 
• Say T. 1823. Descriptions of coleopterous insects collected in the late expedition to the Rocky Mountains, performed by order of Mr. Calhoun, Secretary of War, under Major Long. Journal of the Academy of Natural Sciences of Philadelphia 3: 139–216. 
• Say T. 1825. Descriptions of new species of coleopterous insects inhabiting the United States. Journal of the Academy of Natural Sciences of Philadelphia 5: 160–204. 
• Say T. 1835. Descriptions of new North American insects, and observations on some already described ones. Boston Journal of Natural History 1: 151–203. 
• Schaeffer CW. 1908. On new and known Coleoptera of the families Coccinellidae and Cleridae. Journal of the New York Entomological Society 16(3): 125–135. 
• Schaeffer CW. 1909. New Coleoptera chiefly from Arizona. Bulletin of the Brooklyn Institute of Arts and Sciences 1(15): 375–386. 
• Schenkling S. 1906. Die Cleriden des Deutschen Entomologischen National Museums, nebst Beschreibungen neuer Arten. Deutsche Entomologische Zeitschrift 1: 241–320. 
• Schenkling S. 1910. Cleridae. In: Schenkling S (ed.). Coleoptorum Catalogus, Pars 23. W. Junk; Berlin, Germany. 174 p 
• Solervicens JA. 1986. Revisión taxonómica del género Eurymetopum Blanchard, 1844 (Coleoptera: Cleridae: Phyllobaeninae). Acta Entomológica Chilena 13: 11–120. 
• Spinola MM. 1844. Essai monographique sur les Clérites: Insectes Coléoptères . Volume 2. Imprimiere der frères Ponthenier; Genoa, Italy. 216 p 
• Westcott RL, Merickel FW. 2011. In memoriam. William Frederick Barr (1920-2011). The Coleopterists’ Bulletin 66(2): 100–104. 
• Stocks IC. 2009. Wolcottia (= Isohydnocera) aegra (Coleoptera: Cleridae): association with grasses (Poaceae), native Spartina spp. and the introduced Eragrostis curvula, and resemblance to co-occurring pseudomyrmecine ants. Florida Entomologist 92(4): 648–652. 
• White A. 1849. Nomenclature of coleopterous insects in the collection of the British Museum. Part IV. Cleridae. British Museum (Natural History); London, UK. 68 p 
• Wickham HF. 1895. The Coleoptera of Canada. XIII. The Cleridae of Ontario and Quebec. The Canadian Entomologist 27: 247–253. 
• Wickham HF. 1902. A catalogue of the Coleoptera of Colorado. Bulletin from the Laboratories of Natural History of the State University of Iowa 5(3): 217–310. 
• Wickham HF. 1911. Cleridae. p. 24–25. In: Wickham HF. A list of the Coleoptera of Iowa. Bulletin from the Laboratories of Natural History of the State University of Iowa 35: 1–40. 
• Wickham HF, Wolcott AB. 1912. Notes on Cleridae from North and Central America. Laboratory of Natural History Bulletin, Iowa State University 6(3): 49–67. 
• Wolcott AB. 1908. New species of North American Hydnocera (Coleoptera: Cleridae). The Canadian Entomologist 40: 229–233. 
• Wolcott AB. 1909. The Cleridae of the Public Museum of Milwaukee. The Bulletin of the Wisconsin Natural History Society 7(3–4): 93–102. 
• Wolcott AB. 1910. Cleridae. p. 846–862. In: Blatchley WS. An Illustrated descriptive catalogue of the Coleoptera or beetles known to occur in Indiana. Bulletin of the Indiana Department of Geology and Natural Resources 1: 1–1386. 
• Wolcott AB. 1927. A review of the Cleridae of Costa Rica. Coleopterological Contributions 1(10): 1–104. 
• Wolcott AB. 1928. Descriptions of new species of North American Hydnocerinae (Coleoptera: Cleridae). Entomological News 39: 207–212. 
• Wolcott AB. 1944. A generic review of the subfamily Phyllobaeninae. Journal of the New York Entomological Society 52: 121–152. 
• Wolcott AB. 1947. Catalogue of North American beetles of the family Cleridae. Fieldiana: Zoology 32(2): 59–105. 
• Zeigler D. 1844. Descriptions of new North American Coleoptera. Proceedings of the Academy of Natural Sciences of Philadelphia 2(1844): 43–47.