Abstract

An overview of the genus Psephenops Grouvelle (Coleoptera: Psephenidae: Psepheninae) is pro¬vided with geographic distributions of the species. The taxonomic history of Psephenops smithi Grouvelle, the type species from Grenada and St. Vincent, and junior synonym Xexanchorinus latus Grouvelle from Grenada, is discussed, and lectotypes and paralectotypes are designated for both. Photographs of the type series and a non-type specimen with illustrated male genitalia, along with new distributional information for the species, are provided. The male, female, and associated larva of Psephenops trini Barr and Shepard, new species, are described and illustrated, as is the habitat of this species in Trinidad and Tobago. The Peruvian species Psephenus robacki Spangler is transferred from Psephenus Haldeman to Psephenops as Psephenops robacki (Spangler) new combination. Photographs of the female holotype are included, and possible new distributional records from Peru are reported.

References

• Alkins-Koo M. 1990. The aquatic fauna of two intermittent streams in the southwestern Peninsula, Trinidad. Living World, Journal of the Trinidad and Tobago Field Naturalists’ Club 1989–1990: 36–42.  
• Arce-Pérez R. 2002. A new species of Psephenops Grouvelle (Coleoptera: Psephenidae) from Mexico. Proceedings of the Entomological Society of Washington 104(4): 964–967.  
• Arce-Pérez R, Novelo-Gutiérrez R. 2000. First record of the genus Psephenops (Coleoptera: Psephenidae) from Mexico, with a description of a new species. Entomological News 111(3): 196–200.  
• Arce-Pérez R, Novelo-Gutiérrez R. 2013. Two new species of Psephenops Grouvelle from Belize (Coleoptera: Byrrhoidea: Psephenidae), with a key to the known species from Mexico and Central America. Zootaxa 3670(1): 63–70.  
• Arce-Pérez R, Novelo-Gutiérrez R. 2017. Two new species of Psephenops Grouvelle (Coleoptera: Byrrhoidea: Psephenidae) from Panama. Zootaxa 4323(1): 109–118.  
• Arce-Pérez R, Novelo-Gutiérrez R, Cornejo A. 2015. Psephenops panamaensis sp. nov. (Coleoptera: Byrrhoidea: Psephenidae) from Panama. Zootaxa 4052(2): 233–236.  
• Bameul F. 2001. Un nouveau Psephenops Grouvelle de la Guadeloupe, avec la description de sa larvae (Coleoptera, Psephenidae). Nouvelle Revue d’Entomologie, (New Series) 18(2): 161–172.  
• Bass D. 2000. Freshwater invertebrates. p. 31–33. In: Robinson D, Lowery J (eds.). The natural history of the island of Nevis. Nevis Historical and Conservation Society Press; Ithaca, NY. 69 p 
• Bass D. 2003a. A survey of freshwater macroinvertebrates in Tobago. Living World, Journal of the Trinidad and Tobago Field Naturalists Club 2003: 64–69.  
• Bass D. 2003b. Freshwater macroinvertebrates of Barbados. Journal of the Barbados Museum and Historical Society 49: 269–280.  
• Bass D. 2004a. A survey of freshwater macroinvertebrates on Grenada, West Indies. Living World, Journal of the Trinidad and Tobago Field Naturalists Club 2004: 26–31.  
• Bass D. 2004b. Diurnal stream drift of benthic macroinvertebrates on the small oceanic island of Dominica, West Indies. Caribbean Journal of Science 40: 245–252.  
• Bass D. 2005. A survey of freshwater macroinvertebrates on Antigua, West Indies. Living World, Journal of the Trinidad and Tobago Field Naturalists Club 2005: 11–14.  
• Bass D. 2006. A comparison of the freshwater macroinvertebrate assemblages of St. Kitts and Nevis, West Indies. Living World, Journal of the Trinidad and Tobago Field Naturalists Club 2006: 30–37.  
• Bass D. 2007. Freshwater macroinvertebrates and their habitats in Dominica. Living World, Journal of the Trinidad and Tobago Field Naturalists’ Club 2007: 21–30.  
• Bass D. 2008. Freshwater macroinvertebrates and their habitats in Saba, West Indies. Living World, Journal of the Trinidad and Tobago Field Naturalists Club 2008: 33–38.  
• Bass D. 2009. A comparison of macroinvertebrate communities and their freshwater habitats in the Cayman Islands. Living World, Journal of the Trinidad and Tobago Field Naturalists Club 2009: 1–10.  
• Botosaneanu L, Alkins-Koo M. 1993. The caddis flies (Insecta, Trichoptera) of Trinidad and Tobago, West Indies. Bulletin de l’Institut Royal des Sciences Naturelles de Belgique, Entomologie 63: 5–45.  
• Botosaneanu L, Sakal D. 1992. Ecological observations on the caddisflies (Insecta: Trichoptera) from Trinidad and Tobago (W. Indies). Revue d’hydrobiologie tropicale 25(3): 197–207.  
• Bouysse P, Westercamp D, Andreieff P. 1990. 4. The Lesser Antilles island arc. p. 29–43. In: Moore JC, Mascle A, Taylor E, Underwood MB (eds.). Proceedings of the ocean drilling program, scientific results, vol. 110. College Station, Texas. 448 p 
• Champion GC. 1913. Notes on various Central American Coleoptera, with descriptions of new genera and species. Transactions of the Entomological Society of London 1913(1): 58–169.  
• Darlington PJ Jr. 1936. A list of the West Indian Dryopidae (Coleoptera), with a new genus and eight new species, including one from Colombia. Psyche 43(2–3): 65–83.  
• Delève J. 1967. Contribution a l’etude des Dryopoidea (Coleoptera). XIX. Notes diverses et descriptions d’espèces nouvelles. Bulletin et Annales de la Société Royale Entomologique de Belgique 103: 414–446.  
• Flint OS. 1996. Studies of Neotropical Caddisflies LV: Trichoptera of Trinidad and Tobago. Transactions of the American Entomological Society 122 (2/3): 67–113.  
• Frost CD, Snoke AW. 1989. Tobago, West Indies, a fragment of a Mesozoic oceanic island arc: petrochemical evidence. Journal of the Geological Society 146(6): 953–964.  
• Grouvelle A. 1898. Note V. Clavicornes de Grenada et de St. Vincent (Antilles) récoltés par M. H. H. Smith, et appurtenant au Musée de Cambridge. Notes from the Leyden Museum 20: 35–48.  
• Hinton HE. 1971. The Elmidae (Coleoptera) of Trinidad and Tobago. Bulletin of the British Museum (Natural History), Entomology 26(6): 247–265.  
• Holland WJ. 1919. Herbert Huntingdon Smith. Entomological News 30 (8): 211–214.  
• Hynes HBN. 1948. Notes on the aquatic Hemiptera-Heteroptera of Trinidad and Tobago, B.W.I., with a description of a new species of Martarega B. White (Notonectidae). Transactions of the Royal Entomological Society of London 99(10): 341–360.  
• Lawrence JF, Ślipiński A. 2013. Australian beetles. Volume 1: morphology, classification and keys. CSIRO Publishing; Collingwood, Victoria, Australia. 561 p 
• Lee C-F, Jäch MA, Beutel RG. 2016. 19.7 Psephenidae Lacordaire, 1854. p. 615–628. In: Beutel RG, Leschen RAB (eds.). Coleoptera, Beetles, Volume 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga partim). 684 p. In: Beutel RG, Kristensen NP (eds.) Handbook of Zoology, Arthropoda: Insecta, Second Edition. Walter de Gruyter; Berlin. 
• Lee CF, Sâto M, Shepard WD, Jäch MA. 2007. Phylogeny of Psephenidae (Coleoptera: Byrrhoidea) based on larval, pupal and adult characters. Systematic Entomology 32: 502–538.  
• Meurgey F, Ramage T. 2020. Challenging the Wallacean shortfall: a total assessment of insect diversity on Guadeloupe (French West Indies), a checklist and bibliography. Insecta Mundi 786: 1–183.  
• Michalski J. 1988. A catalog and guide to the dragonflies of Trinidad (Order Odonata). Zoology Department, University of the West Indies, Trinidad, Occasional Paper No. 6: 1–146.  
• Neiser N, Alkins-Koo M. 1991. The water bugs of Trinidad and Tobago. Zoology Department, University of the West Indies, Trinidad, Occasional Paper No. 9: 1–127.  
• Pararas-Carayannis G. 2010. Assessment of the tsunamigenic potential along the northern Caribbean margin – Case study: earthquake and tsunamis of 12 January 2010 in Haiti. Science of Tsunami Hazards 29(3): 175–201.  
• Peck SB. 2006. The beetle fauna of Dominica, Lesser Antilles (Insecta: Coleoptera): diversity and distribution. Insecta Mundi 20 (3–4): 165–209.  
• Peck SB. 2009a. The beetles of Barbados, West Indies (Insecta: Coleoptera): diversity, distribution and faunal structure. Insecta Mundi 73: 1–51.  
• Peck SB. 2009b. The beetles of St. Lucia, Lesser Antilles (Insecta: Coleoptera); diversity and distributions. Insecta Mundi 106: 1–34.  
• Peck SB. 2010. The beetles of the island of St. Vincent, Lesser Antilles (Insecta: Coleoptera); diversity and distributions. Insecta Mundi 144: 1–77.  
• Peck SB. 2011a. The diversity and distributions of the beetles (Insecta: Coleoptera) of the northern Leeward Islands, Lesser Antilles (Anguilla, Antigua, Barbuda, Nevis, Saba, St. Barthélemy, St. Eustatius, St. Kitts and St. Martin-St. Maarten. Insecta Mundi 159: 1–54.  
• Peck SB. 2011b. The beetles of Martinique, Lesser Antilles (Insecta: Coleoptera); diversity and distributions. Insecta Mundi 178: 1–57.  
• Peck SB. 2016. The beetles of the Lesser Antilles (Insecta: Coleoptera): diversity and distributions. Insecta Mundi 460: 1–360.  
• Peck SB, Cook J, Hardy JD Jr. 2002. Beetle fauna of the island of Tobago, Trinidad and Tobago, West Indies. Insecta Mundi 16(1–3): 9–23.  
• Peck SB, Thomas MC, Turnbow RH Jr. 2014. The diversity and distributions of the beetles (Insecta: Coleoptera) of the Guadeloupe Archipelago (Grande-Terre, Basse-Terre, La Désirade, Marie-Galante, Le Saintes, and Petite-Terre), Lesser Antilles. Insecta Mundi 352: 1–156.  
• Shepard WD, Barr CB. 2023. A revision of the Chilean water penny genus Tychepsephus Waterhouse, 1876 (Coleoptera, Psephenidae, Eubriinae), with description of a second species and two larval morphotypes, and notes on other Chilean Psephenidae. ZooKeys 1164: 23–61.  
• Shorthouse DP. 2010. SimpleMappr, an online tool to produce publication-quality point maps. Available at https://www.simplemappr.net  (Last accessed 1 December 2023.)
• Spangler PJ. 1966. The Catherwood Foundation Peruvian-Amazon Expedition. XIII—The aquatic Coleoptera (Dytiscidae; Noteridae; Gyrinidae; Hydrophilidae; Dascillidae; Helodidae; Psephenidae; Elmidae). p. 377–443. In: Patrick R (ed.). The Catherwood Foundation Peruvian-Amazon Expedition: limnological and systematic studies. Monographs of the Academy of Natural Sciences of Philadelphia 14: 1–495.  
• Spangler PJ. 1990. A new species and new record of the water-penny genus Psephenops (Coleoptera: Psephenidae) from Costa Rica. Entomological News 101(3): 137–140.  
• Speed RC. 1981. Geology of Barbados: Implications for an accretionary origin. Oceanologica Acta 1981: 259–265.  
• Stark BP. 1994. Anacroneuria of Trinidad and Tobago (Plecoptera: Perlidae). Aquatic Insects 16(3): 171–175.