Historical and morphological review of the subgenus Indonthophagus Kabakov, 2006 of Onthophagus Latreille, 1802 (Coleoptera: Scarabaeidae: Scarabaeinae: Onthophagini)
Stefano Ziani
GeoL@b APS, Via Laderchi, 3, 48018 Faenza (RA), Italy
Abstract
The subgenus Indonthophagus Kabakov, 2006 of Onthophagus Latreille, 1802 (Coleoptera: Scara¬baeidae: Scarabaeinae: Onthophagini) is reviewed. A morphological definition of it is provided. A diagnosis, based both on external and internal characters, of the seven species deemed to belong to the group according to literature, is supplied. As a result, Onthophagus spinifex (Fabricius, 1781), previously placed with some uncertainty in the subgenus, is definitively excluded. A key to Indonthophagus species is also provided, as well as images of male, aedeagus and endophallus for each species. New country records of Onthophagus turbatus Walker, 1858 for Pakistan, Nepal and Bhutan are given.
References
- Alfieri A. 1976. The Coleoptera of Egypt. Mémoires de la Société entomologique d’Égypte 5: 1–287.
- Ambrus R, Grosser W. 2012. Contribution to the knowledge of longhorn beetles from Dhofar region in sultanate of Oman (Coleoptera: Cerambycidae). Humanity Space, International Almanac 1(2): 448–457.
- Anu A, Vinod KV. 2017. Succession of Dung Beetles (Scarabaeinae: Coleoptera) in the Cow Dung Pats of a Tropical Agro habitat in Kerala, South India. Journal of Environmental Science, Toxicology and Food Technology 11(6): 46–49. https://doi.org/10.9790/2402-1106024649
- Arrow GJ. 1907. Some new species and genera of Lamellicorn Coleoptera from the Indian Empire (part II). The Annals and Magazine of natural History, including Zoology, Botany and Geology 7 (19): 416–439. https://doi.org/10.1080/00222930708562664
- Arrow GJ. 1931. The fauna of British India, including Ceylon and Burma. Coleoptera Lamellicornia. Part III (Coprinae). Taylor and Francis; London. 428 p
- Asha G, Manoj K, Megha PP, Sinu PA. 2021. Spatiotemporal effects on dung beetle activities in island forests-home garden matrix in a tropical village landscape. Scientific Reports 11: 1–3. https://doi.org/10.1038/s41598-021-96831-5
- Asha G, Manoj K, Rajesh TP, Varma A, Ballullaya UP, Sinu PA. 2022. Dung beetles prefer used land over natural greenspace in urban landscape. Scientific Reports 12(1): 1. https://doi.org/10.1038/s41598-022-26841-4
- Aston P, Vor Y. 2008. Dung beetles of Hong Kong. Part 2. Hong Kong Discovery 46: 59–63.
- Balthasar V. 1935. Onthophagus–Arten Chinas, Japans und der angrenzenden Länder. Mit Beschreibungen von 14 neuen Arten und einer Unterart (30. Beitrag zur Kenntnis der palaearktischen Scarabaeiden). Folia Zoologica et Hydrobiologica 8 (2): 303–358.
- Balthasar V. 1942. Die Coprophagen der chinesischen Provinz Fukien (70. Beitrag zur Kenntnis der Scarabaeiden). Entomologische Blätter 38(4): 113–125.
- Balthasar V. 1956. Scarabaeidae (Coleoptera) der Afghanistan–expedition (1952 u. 1953) J. Klapperichs. Scarabaeidae (Coleoptera) z expedice J. Klapperichia do Afghanistanu v letech 1952 a 1953. Acta entomologica Musei Nationalis Pragae 30 [1955]: 409–439.
- Balthasar V. 1963. Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Coleoptera Lamellicornia. Band 2. Coprinae (Onitini, Oniticellini, Onthophagini). Tschechoslowakischen Akademie der Wissenschaften; Prague. 627 p. + 16 pls
- Balthasar V. 1968. Beiträge zur Kenntnis der Fauna Afghanistans (Sammelergebnisse von O. Jakeš 1963–64, D. Povolný 1965, D. Povolný & Fr. Tenora 1966, J. Šimek 1965–66, D. Povolný, I. Geisler, Z. Šebek & Fr. Tenora 1967). Lamellicornia Col. (Nachtrag). Acta Musei Moraviae Supplementum 53: 147–156.
- Balthasar V. 1969. Contributions à la connaissance de la faune entomologique de la Côte-d’Ivoire (J. Decelle, 1961–1964). XXXII. Coleoptera Scarabaeidae et Aphodiidae. Annales du Musée Royal de l’Afrique Centrale/Sciences Zoologiques 175: 59–73.
- Balthasar V. 1972. Coleoptera aus Nordostafrika. Scarabaeoidea: Scarabaeidae und Aphodiidae. 135. Beitrag zur Kenntnis der Scarabaeoidea. Notulae Entomologicae 52: 18–32.
- Baraud J. 1968. Contribution à la faune de l’Iran. 10. Coléoptères Scarabaeoidea. Annales de la Société Entomologique de France (N. S.) 4(4): 915–925. https://doi.org/10.1080/21686351.1968.12277995
- Baraud J. 1985. Coléoptères Scarabaeoidea, Faune du Nord de l’Afrique, du Maroc au Sinaï. Éditions Lechevalier; Paris. 650 p
- Barbero E, Borghesio L, Dellacasa G, Dellacasa M. 1998. Contribution to the knowledge of Scarabaeidae and Aphodiidae from southern Ethiopia (Coleoptera Scarabaeoidea). Bollettino della Società Entomologica Italiana 130(3): 233–254.
- Binaghi G, Dellacasa G, Poggi R. 1969. Nuovi caratteri diagnostici per la determinazione degli Onthophagus del gruppo ovatus (L.) e geonemia controllata delle specie italiane del gruppo. Memorie della Società Entomologica Italiana 48: 29–46.
- Biswas S, Chatterjee SK. 1985. Insecta: Coleoptera: Scarabaeidae: Coprinae. Records of the Zoological Survey of India 82(1–4): 147–177. https://doi.org/10.26515/rzsi/v82/i1-4/1984/161291
- Biswas S, Chatterjee SK. 1986a. Dung beetles fauna (Coleoptera: Scarabaeidae: Scarabaeinae) of Palamou Tiger Reserve, Bihar, with description of a new species. Records of the Zoological Survey of India 84(3–4): 57–67. https://doi.org/10.26515/rzsi/v83/i3-4/1986/161311
- Biswas S, Chatterjee SK. 1986b. Scarabaeidae (India: Coleoptera) of Silent Valley, Kerala, India with descriptions of three new species. Records of the Zoological Survey of India 84(1–4): 79–96. https://doi.org/10.26515/rzsi/v84/i1-4/1986/161083
- Biswas S, Ghosh AK. 2000. Coleoptera: Scarabaeidae: Scarabaeinae. p. 513–623. In: Alfred JRB (ed.). Fauna of Meghalaya: Part 5 (Insects). Zoological Survey of India; Kolkata. 666 p
- Biswas S, Mulay SV. 2001. Insecta: Scarabid (Coleoptera). p. 129–142. In: Cherian PT (ed.). Fauna of Nilgiri Biosphere Reserve, fauna of conservation areas series 11. Zoological Survey of India; Kolkata. 330 p
- Blanchard CE. 1847. Insectes de l’Amérique méridionale. Recueillis par Alcide d’Orbigny et décrits par Emile Blanchard et Auguste Brullé. p. 57–222. In: Voyage dans l’Amérique méridionale (le Brésil, la République orientale de l’Uruguay, la République Argentine, la Patagonie, la République du Chili, la République de Bolivie, la République du Pérou), exécuté pendant les années 1826, 1827, 1828, 1829, 1830, 1831, 1832 et 1833 par Alcide d’Orbigny. Ouvrage dédié au Roi, et publié sous les auspices de M. le Ministre de l’Instruction publique (commencé sous le ministère de M. Guizot). Tome sixième. 2.e Partie: Insectes. P. Bertrand; Paris [&] V. Levrault; Strasbourg. 4 + 222 p.
- Boheman CH. 1860. Coleoptera samlade af J. A. Wahlberg i sydvestra Afrika. Kungliga Svenska vetenskapsakademiens handlingar 17 (3): 107–120.
- Bouchard P, Bousquet Y. 2020. Additions and corrections to ‘Family–group names in Coleoptera (Insecta)’. Zookeys 922: 65–139. https://doi.org/10.3897/zookeys.922.46367
- Boucomont A. 1914. Onthophagus asiatiques nouveaux ou peu connus. Annali del Museo Civico di Storia Naturale di Genova 46(= ser. 3, 6): 210–243. https://doi.org/10.5962/bhl.part.14786
- Boucomont A. 1927. Zur Erforschung des Persischen Golfes. (Beitrag Nr. 6). Entomologische Mitteilungen 16 (3): 187–188.
- Boucomont A. 1929. A list of the coprophagous Coleoptera of China. Lingnan Science Journal 7: 759–794.
- Boucomont A, Gillet G. 1921. Faune entomologique de l’Indochine française. Fam. Scarabaeidae Laparosticti (Coléoptères). Imprimerie Nouvelle Albert Portail 4: 1–76.
- Boucomont A, Gillet JJE. 1927. Fam. Scarabaeidae. Subfam. Coprinae II. p. 103–263. In: Gillet JJE, Boucomont A (eds.). Scarabaeidae: Coprinae, Termitotroginae. Partes 38 et 90, Vol. 19 (3). In: Schenkling S (ed.). Coleopterorum Catalogus. W. Junk; Berlin. 265 p
- Bousquet Y. 2016. Litteratura Coleopterologica (1758–1900): a guide to selected books related to the taxonomy of Coleoptera with publication dates and notes. ZooKeys 583: 1. https://doi.org/10.3897/zookeys.583.7084
- Bunalski M, Samin N, Ghahari H, Hawkeswood TJ. 2014. Contributions to the knowledge the scarab beetles of Golestan province, Northern Iran with checklist of Iranian Scarabaeoidea (Coleoptera). Polish Journal of Entomology 83: 141–170. https://doi.org/10.2478/pjen-2014-0011
- Burmeister HCC. 1846. Genera quaedam insectorum. Iconibus illustravit et descripsit. Volumen I. Continet tabulas XL. Heft 10. A. Burmeister; Berlin. viii + [2] + [120] p
- Carpaneto GM, Piattella E. 1990. Analisi zoogeografica preliminare dei Coleotteri Scarabeidi della Somalia (Coleoptera, Scarabaeidae s. str.). Biogeographia 14(1988): 265–292. https://doi.org/10.21426/B614110347
- B614110347 Chandra K. 2005. Insecta: Coleoptera: Scarabaeidae. p. 141–155. In: Metha HS (ed.). Fauna of Western Himalaya (Part–2). Himachal Pradesh. Zoological Survey of India; Kolkata. 359 p
- Chandra K, Gupta D. 2011. Two new records of genus Onthophagus Latreille, 1802 (Coleoptera: Scarabaeidae) from Madhya Pradesh, India. Uttar Pradesh Journal of Zoology 31(2): 253–255.
- Chandra K, Gupta D. 2012. First report on five species of genus Onthophagus Latreille, 1802 (Coleoptera, Scarabaeidae) from Madhya Pradesh, India, and their description of external male genitalia. Biodiversity Journal 3(2): 99–106.
- Chandra K, Gupta D. 2013a. Scarab beetles (Coleoptera: Scarabaeoidea) of Barnawapara Wildlife sanctuary, Chhattisgarh, India. Journal of Threatened Taxa 5(12): 4660–4671. https://doi.org/10.11609/JoTT.o3251.4660-71
- Chandra K, Gupta D. 2013b. Taxonomic studies on dung beetles (Coleoptera: Scarabaeidae, Geotrupidae, Hybosoridae) of Chhattisgarh, India. Munis Entomology & Zoology 8(1): 331–360.
- Chandra K, Gupta D, Uniyal VP, Bharadwaj M, Sanyal AK. 2012. Studies on Scarabaeid beetles (Coleoptera) of Govind Wildlife Sanctuary, Gharwal, Uttarakhand, India. Biological Forum 4(1): 48–54.
- Chen J. 2002. Dung beetle of the world – An introduction to Scarabaeinae of the world. Owl Press; Taipei. 303 p
- Cheung JKH, Wong PMW, Leung MH, Lee JM. 2020. Scarabaeinae in Hong Kong: The first record of tribe Deltochilini, genus Panelus and three species (Panelus parvulus, Panelus tonkinensis and Onthophagus vigilans), and the updated species checklist. Hong Kong Biodiversity 26: 25–28.
- Cheung K, Bai M, Leung MH, Lee Y, Chan K, Yang X. 2018. Scarabaeinae (Coleoptera: Scarabaeidae) from Hong Kong. Zoological Systematics 43(3): 233–267.
- Dejean PFMA. 1821. Catalogue de la collection de coléoptères de M. le Baron Dejean. Crevot; Paris. 136 p https://doi.org/10.5962/bhl.title.11259
- Dejean PFMA. 1833. Catalogue des Coléoptères de la collection de M. le Comte Dejean. Fascicle 2: 97–176. https://doi.org/10.5962/bhl.title.8771
- Dejean PFMA. 1836. Catalogue des Coléoptères de la collection de M. le Comte Dejean. Troisième édition, revue, corrigée et augmentée. Fascicles 1–4: 97–176. https://doi.org/10.5962/t.173109
- Dellacasa G, Bordat P, Dellacasa M. 2001. A revisional essay of world genus-group taxa of Aphodiinae (Coleoptera Aphodiidae). Memorie della Società Entomologica Italiana 79: 1–482.
- Dohrn CA. 1868. Eine Todtenkranz. Bernt Wilhelm Westermann. Entomologische Zeitung 29(1–3): 215–218.
- Orbigny H. 1898. Synopsis des Onthophagides paléarctiques. L’Abeille 29: 117–254.
- Orbigny H. 1900. Supplément au synopsis des Onthophagides paléarctiques. L’Abeille 29: 289–300.
- Orbigny H. 1902. Mémoire sur les Onthophagides d’Afrique. Annales de la Société Entomologique de France 71(1–2): 1–324.
- Orbigny H. 1904. Onthophagides africains de la collection du Musée Civique de Gênes. Annali del Museo Civico di Storia Naturale di Genova 41(= ser. 3, 1): 253–331.
- Orbigny H. 1905. Onthophagides provenant du voyage de M. Alluaud dans l’Afrique orientale et descriptions d’autres espèces africaines nouvelles. Annales de la Société Entomologique de France 74(3–4): 381–536.
- Orbigny H. 1908. Descriptions d’espèces nouvelles d’Onthophagides africains et notes synonymiques. Annales de la Société Entomologique de France 77: 65–208. https://doi.org/10.5962/bhl.part.5618
- Orbigny H. 1913. Synopsis des Onthophagides d’Afrique. Annales de la Société Entomologique de France 82: 1–608.
- Orbigny H. 1922. Coléoptères Onthophagines. p. 599–615. In: Rothschild MD (ed.). Voyage de M. le Baron Maurice de Rotschild en Éthiopie et en Afrique Orientale Anglaise (1904–1905). Résultats scientifiques. Animaux Articulés. Imprimerie Nationale; Paris. 1041 p
- El–Zoheiry MS, Mohamed N. 1949. List of Egyptian insects in the collection of the Entomological Section. Ministry of Agricolture, Government Press; Cairo. 87 p
- Encyclopaedia Britannica. 2018. "East Indies". Encyclopaedia Britannica, 1 Jan. 2018 Available at https://www.britannica.com/place/East-Indies (Last accessed November 2022.)
- Endrődi S. 1974. The scientific results of Dr. Gy. Topál’s collecting in India. No. 5. Sammelergebnisse aus der Superfamilie Lamellicornia (Coleoptera). Acta Musei Silesiae Series A 23: 1–5.
- Fabricius JC. 1781. Species insectorum exhibentes eorum differentias specificas, synonyma auctorum, loca natalia, metamorphosin adiectis observationibus, descriptionibus. Tom. I. Carol. Ernest; Bonn, Hamburg and Kiel. viii + 552 p https://doi.org/10.5962/bhl.title.36509
- Fabricius JC. 1787. Mantissa insectorum sistens eorum species nuper detectas adiectis characteribus genericis, differentiis specificis, emendationibus, observationibus. Tom. I. Christ. Gottl. Proft; Copenhagen. xx + 348 p. https://doi. org/10.5962/bhl.title.11657 https://doi.org/10.5962/bhl.title.11657
- Fabricius JC. 1792. Entomologia systematica emendata et aucta. Secundum classes, ordines, genera, species adjectis synonimis, locis, observationibus, descriptionibus. Tom. I. [Pars I]. Christ. Gottl. Proft; Copenhagen. xx + 330 p. https://doi. org/10.5962/bhl.title.122153 https://doi.org/10.5962/bhl.title.122153
- Fabricius JC. 1801. Systema eleutheratorum secundum ordines, genera, species: adiectis synonymis, locis, observationibus, descriptionibus. Tomus I. Bibliopolii Academici Novi; Kiel. xxiv + 506 p
- Fairmaire L. 1887. Coléoptères de l intérieur de la Chine. Annales de la Société Entomologique de Belgique 31: 87–136.
- Ferreira MC. 1972. Os Escarabídeos de África (sul do Sáara). I. Revista de Entomologia de Moçambique 11[1968–1969]: 5–1088.
- Frey G. 1956. Liste der von S. Nathan in der Wüste Thar im Jahre 1955 gesammelten Lamellicorniern mit der Beschreibung einer neuen Catharsius–Art (Col.). Entomologische Arbeiten aus dem Museum G. Frey 7: 654–656.
- Gajendra N, Prasad SK. 2016. A review of coleoptera diversity of Chhattisgarh: Updated checklist 2015. International Journal of Science and Research 5(4): 710–714. https://doi.org/10.21275/v5i4.NOV162650
- Génier F. 2019. Endophallites: a proposed neologism for naming the sclerotized elements of the insect endophallus (Arthropoda: Insecta). Annales de la Société entomologique de France (N.S.) 55(6): 482–484. https://doi.org/10.1080/00379271.2019.1685907
- Génier F, Moretto P. 2017. Digitonthophagus Balthasar, 1959: taxonomy, systematics, and morphological phylogeny of the genus revealing an African species complex (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 4248: 1–110. https://doi.org/10.11646/zootaxa.4248.1.1
- Germar EF. 1813. VI. Neue Insekten, beschrieben vom Herausgeber. Magazin der Entomologie 1: 114–133.
- Gestro R. 1889. Viaggio ad Assab nel Mar Rosso dei Signori G. Doria ed O. Beccari con il R. Avviso “Esploratore” dal 16 Novembre 1879 al 26 Febbraio 1880. IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova 27(= ser. 2, 7): 5–72.
- Ghosh J, Das P, Ghosh SK, Bhunia D, Kushwaha RK, Gupta D, Chandra K. 2020. Insecta: Coleoptera. Fauna of Haryana, State Fauna Series 24(1): 221–275.
- Ghosh J, Gupta D, Chandra K, Das P, Bhunia D. 2022a. Insecta: Coleoptera: Scarabaeoidea. p. 1–28. In: Sidhu AK, Pathania PC, Gupta D (eds.). Fauna of Himachal Pradesh. State Fauna Series 26 (2 [2021]). Zoological Survey of India; Kolkata. 528 p
- Ghosh J, Gupta D, Chandra K, Saha GK. 2022b. Scarab beetles (Coleoptera: Scarabaeidae) of North-East India. Nature Books India; New Delhi. 200 p
- Ghosh J, Saha GK, Gupta D, Chandra K. 2021. Checklist of scarab beetles (Coleoptera: Scarabaeidae) from Tripura, India. Records of the Zoological Survey of India 121(3): 383–391.
- Gillet JJE. 1924. Descriptions d’Onthophagus nouveaux d’Asie tropicale. Annales de la Société scientifique de Bruxelles 44: 66–70.
- Gillett MPT. 1995. Scarab dung beetles in the Al Ain Region of the UAE and neighbouring areas of Oman (Coleoptera: Scarabaeidae). Tribulus 5(1): 14–23.
- Gillett MPT, Gillett CPDT. 2005. Insects & other arthropods. p. 168–194. Species checklist. p. 352–356. In: Aspinall S, Hellyer P (eds.). The Emirates. A natural history. P. Vine, Trident Press; London. 428 p
- Gmelin JF. 1790. Caroli a Linné Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio decima tertia, aucta, reformata. Tom. I. Pars IV. Georg Emanuel Beer; Leipzig. 1517–2224 p
- Gupta AK. 1989. Studies on spermathecal capsules of some Scarabaeinae beetles (Coleoptera: Scarabaeidae) of North–West India. Y.E.S. Quarterly 6(3): 22–28.
- Gupta AK, Mittal IC. 1987. Studies on the male genitalia of some Indian dung beetles (Coleoptera; Scarabaeidae; Scarabaeinae) and their Taxonomic Significance. Y.E.S. Quarterly 4 (3): 46–58.
- Gupta D, Chandra K, Khan S. 2014. An updated checklist of scarabaeoid beetles (Coleoptera: Scarabaeoidea) of Pench Tiger Reserve, Madhya Pradesh, India. Journal of Entomology and Zoology Studies 2(5): 225–240.
- Gupta D, Chandra K, Das P, Ghosh J. 2018. Chapter 35. Insecta: Coleoptera. p. 399–590. In: Chandra K, Gupta D, Gopi KC, Tripathy B, Kumar V (eds.). Faunal diversity of Indian Himalaya. Zoological Survey of India; Kolkata. 872 p
- Gupta D, Ghosh J, Dutta S, Das P, Chandra K. 2022. Chapter 45. Insecta: Coleoptera. p. 407–461. In: Banerjee D, Chandra K, Raghunathan C, Singh N, Gupta D (eds.). Faunal diversity of biogeographic zones of India: Deccan Peninsula. Zoological Survey of India; Kolkata. 647 p
- Hanboonsong Y, Chunram S, Pimpasalee S, Emberson RW, Masumoto K. 1999. The dung beetle fauna (Coleoptera, Scarabaeidae) of Northeast Thailand. Elytra 27(2): 463–469.
- Harold E von. 1859. Beiträge zur Kenntniss einiger coprophagen Lamellicornien. (Erstes Stück). Berliner Entomologische Zeitschrift 3: 193–224. https://doi.org/10.1002/mmnd.18590030208
- Harold E von. 1862. Beiträge zur Kenntniss einiger coprophagen Lamellicornien (Viertes Stück). Berliner Entomologische Zeitschrift 6: 379–403. https://doi.org/10.1002/mmnd.47918620310
- Harold E von. 1867. III. Beitrage zur Kenntniss der Gattung Onthophagus vom Herausgeber. I. Theil: Australier und Afrikaner. Coleopterologische Hefte 2: 23–59.
- Harold E von. 1869. Tom. IV. Scarabaeidae. p. 979–1346. In: Gemminger M, Harold E von (eds.). Catalogus coleopterorum hucusque descriptorum synonymicus et systematicus. E. H. Gummi; Munich. 3822 p
- Harold E von. 1870. Berichtigungen und Zusätze zum Catalogus Coleopterorum synonymicus et systematicus. 2. Vom Herausgeber. Ad Vol. IV. Coleopterologische Hefte 6: 104–110.
- Harold E von. 1872. Berichtigungen und Zusätze zum Catalogus Coleopterorum synonymicus et systematicus. 2. Vom Herausgeber. Coleopterologische Hefte 9–10: 205–207.
- Harold E von. 1879. II. Diagnosen neuer Arten. Coleopterologische Hefte 16: 225–232.
- Harold E von. 1880. Einige neue Coleopteren. Mitteilungen des Münchener Entomologischen Vereins 4: 148.
- Hashmi AA, Tashfeen A. 1992. Coleoptera of Pakistan. Proceedings of Pakistan Congress of Zoology 12: 133–170.
- Herbst JFW. 1786. Erste Mantisse zum Verzeichniss der ersten Klasse meiner Insectensammlung. Archiv der Insectengeschichte. Herausgegeben von Johann Caspar Füessly 7–8: 153–182.
- Hope FW. 1837. The coleopterist’s manual, containing the lamellicorn insects of Linnaeus and Fabricius. Henry G. Bohn; London. 121 p.
- Hope FW. 1838. Observations on the Lamellicorns of Olivier. Entomological Magazine 5: 312–326.
- Howden HF, Cartwright OL. 1963. Scarab beetles of the genus Onthophagus Latreille North of Mexico (Coleoptera: Scarabaeidae). Proceedings of the United States National Museum 114: 1–135. https://doi.org/10.5479/si.00963801.114-3467.1
- Hua L–z. 2002. List of Chinese insects. Vol. II. Zhongshan (Sun Yat–sen) University Press; Guangzhou. 612 p
- Hwang K. 1952. List of recorded species of Chinese Scarabaeidae (A supplement to Wu’s Catalogus Insectorum Sinensium). Acta Entomologica Sinica 2(2): 136–154.
- ICZN [International Commission of Zoological Nomenclature]. 1993. Opinion 1710. J.C. Megerle’s (1801–1805) auction catalogues of insects: suppressed for nomenclatural purposes, with the specific names of Saperda alboguttata Megerle, 1803 (currently Apomecyna alboguttata; Coleoptera) and Hippobosca variegata Megerle, 1803 (Diptera) conserved. Bulletin of Zoological Nomenclature 50: 79–82.
- ICZN [International Commission of Zoological Nomenclature]. 1999. International Code of Zoological Nomenclature (4th ed.). International Trust for Zoological Nomenclature; London. 306 p
- Illiger JCW. 1800. Olivier’s Entomologie oder Naturgeschichte der Insekten mit ihren Gattungs-und ArtMerkmalen, ihrer Beschreibung und Synonymie. Käfer. Uebersetzt und mit Zusätzen und Anmerkungen durchgängig begleitet von Karl Illiger. Erster Theil. Mit Kupfern. Karl Reichard; Braunschweig. xvi + 309 p
- Illiger JCW. 1802. Zusätze, Berichtigungen und Bemerkungen zu Fabricii Systema Eleutheratorum. Magazin für Insektenkunde 1: 306–536.
- Jablonsky CG, Herbst JFW. 1789. Natursystem aller bekannten in- und ausländischen Insekten, als eine Fortsetzung der von Büffonschen Naturgeschichte. Nach dem System des Ritters von Linné und Fabricius zu bearbeiten angefangen. Der Käfer zwenter Theil. Mit siebzehn illuminirten Kupfertafeln. Hefte 2. Joachim Pauli; Berlin. xvi + lxiv + 330 p
- Jadhav MJ, Sharma RM. 2012. Insecta: Coleoptera: Scarabaeidae (Scarabaeid Beetles). p. 489–494. In: Venkataraman K (ed.). Fauna of Maharashtra 2, State Fauna Series 20. Zoological Survey of India; Kolkata. 528 p
- Janíková M. 1998. Coleoptera. Beetles checklist. Al Nahda Printing Press; Muscat. 39 p
- Kabakov ON. 2006. Plastinchatousye zhuki podsemeystva Scarabaeinae (Insecta: Coleoptera: Scarabaeidae) fauny Rossii i sopredel’nykh stran. Tovarishchestvo nauchnykh izdanii KMK; Moscow. 374 p
- Kabakov ON, Napolov A. 1999. Fauna and ecology of Lamellicornia of subfamily Scarabaeinae (Coleoptera, Scarabaeidae) of Vietnam and some parts of adjacent countries: South China, Laos and Thailand. Latvijas Entomologs 37: 58–96.
- Kabakov ON, Yanushev VV. 1983. Materialy po faunistike i ekologii zhukov roda Onthophagus (Scarabaeidae) iz yugo– vostochnoy Azii. p. 156–165. In: Medvedev LN (ed.). Fauna i ekologiyazhivotnykh Vietnama. Akademiya Nauk SSSR; Moscow. 207 p
- Kalawate AS, Mukhopadhyay B, Pawar SV, Shinde VD. 2021. Some new records of scarab beetles of the genus Onthophagus Latreille, 1802 (Coleoptera: Scarabaeidae) from northern Western Ghats, Maharashtra, with a checklist. Journal of Threatened Taxa 13(1): 17580–17586. https://doi.org/10.11609/jott.5695.13.1.17580-17586
- Karimbumkara SN, Rajan PD. 2013. Insecta Coleoptera: Scarabaeidae Scarabaeinae Dung Beetles. p. 173–178. In: Director of the Zoological Survey of India (ed.). Fauna of Karnataka, State Fauna Series. Zoologica Survey of India; Kolkata. 595 p
- Kaur P, Yadav AS. 2014. New cytological data of dung beetle species from the genus Onthophagus Latreille (Coleoptera: Scarabaeidae, Scarabaeinae) from Haryana. Journal of Entomology and Zoology Studies 2(3): 1–6.
- Kaur P, Yadav AS. 2018. Cytogenetic analysis, karyotype evolution and phylogenetic study of family Scarabaeidae (Coleoptera: Insecta). Research Review 3(8): 390.
- Kharel BP, Schoolmeesters P, Sarkar SK. 2020. A first faunistic account on the Onthophagus Latreille, 1802 (Coleoptera, Scarabaeidae, Scarabaeinae) of the Nadia district, West Bengal, with a preliminary checklist from India. Check List 16(2): 361–381. https://doi.org/10.15560/16.2.361
- Klug JCF. 1845. Symbolae physicae, seu icones et descriptiones insectorum, quae ex itinere per Africam borealem et Asiam occidentalem Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg studio novae aut illustratae redierunt. Precensuit Dr. Fr. Klug regis iussu et impensis eddit Dr. C.G. Ehrenberg. Decas quinta. Mittler; Berlin. (41) p. + pls. 41–50.
- Klug JCF. 1855. Diagnosen der neuer Coleopteren aus Mossambique. Berichten der Königlich-Preussischen Akademie der Wissenschaften 20: 643–660.
- Klug JCF. 1862. Coleoptera. Käfer. p. 145–267 + pls 8–15. Naturwissenschaftliche Reise nach Mossambique auf Befehl seiner Majestät des Königs Friedrich Wilhelm IV in den Jahren 1842 bis 1848 ausgeführt von Wilhelm C.H. Peters. Zoologie. V. Insecten und Myriopoden. Bearbeitet in Verbindung mit Klug, Loew, Schaum, Hagen, Gerstaecker, Hopffer. Mit fünf und dreissig Kupfertafeln.Georg Reimer; Berlin. xxi + 566 p
- Kolenati FA. 1846. Meletemata entomologica [Insecta Caucasi. Coleoptera, Dermaptera, Lepidoptera, Neuroptera, Mutillidae, Aphaniptera, Anoplura]. Fasc. V. Accedunt tabulae III. Imperialis Academiae Scientarium; St. Petersburg. (4) + iii + 169 + [1] p. + pls 17–19
- Krajcik M. 2006. Checklist of Scarabaeoidea of the world. 1. Scarabaeinae (Coleoptera: Scarabaeidae: Scarabaeinae). Animma.x 3, supplement: 1–189.
- Krajcik M. 2012. Checklist of the world Scarabaeoidea. Animma.x 5, supplement: 1–278.
- Krajcik M. 2013. Addenda to checklist of world Scarabaeoidea 2012. Animma.x 49–54: 1–299. https://doi.org/10.3233/BIR-2012-0614
- Král D, Batelka J. 2017. Order Coleoptera, superfamily Scarabaeoidea. Arthropod Fauna of the UAE 6: 78–168.
- Lansberge JW van. 1885. Descriptions d’espèces nouvelles de Coléoptères appartenant au Musée civique de Gênes. Annali del Museo Civico di Storia Naturale di Genova 22(= ser. 2, 1): 375–400.
- Latha T. 2019. Seasonal activity of dung beetles (Scarabaeinae) in a forest in South Western Ghats. International Journal of Forest, Animal and Fisheries Research 3(2): 58–64. https://doi.org/10.22161/ijfaf.3.2.2
- Latha T, Thomas SK. 2018a. Species list with pictorial key for dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) of Nelliampathi in South Western Ghats, India. International Journal of Current Advanced Research 7(10): 16121–16128.
- Latha T, Thomas SK. 2018b. Dung beetle (Coleoptera: Scarabaeinae) community structure across a forest-agriculture habitat ecotone in South Western Ghats. International Journal of Environment, Agriculture and Biotechnology 3(5): 1879. https://doi.org/10.22161/ijeab/3.5.41
- Latreille PA. 1802. Histoire Naturelle, Générale et Particulière, des Crustacés et des Insectes; Ouvrage faisant suite à l’Histoire Naturelle, générale et particulière, composée par Leclerc de Buffon, et rédigée par C. S. Sonnini, membre de plusieurs Sociétés savantes. Famille Naturelle des Genres. Tome Troisième. F. Dufart; Paris. xii + 464 p https://doi.org/10.5962/bhl.title.15764
- Lau CSK. 2019. Checklist of Insects of Hong Kong. Agriculture, Fisheries and Conservation Department; Hong Kong. 1058 p
- Löbl I, Krell F-T, Ziani S, Král D. 2006. Scarabaeidae, subfamily Scarabaeinae, tribe Onthophagini. p. 159–176. In: Löbl I, Smetana A (eds.). Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea – Scirtoidea – Dascilloidea – Buprestoidea – Byrrhoidea. Apollo Books; Stenstrup. 690 p
- Löbl I, Löbl D. 2016. Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea – Scirtoidea – Dascilloidea – Buprestoidea – Byrrhoidea. Revised and updated edition. Brill; Leiden, Boston. 983 p https://doi.org/10.1163/9789004309142
- Löbl I, Smetana A. 2006. Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea – Scirtoidea – Dascilloidea – Buprestoidea – Byrrhoidea. Apollo Books; Stenstrup. 690 p
- Marseul SA. 1866. Catalogue des Coléoptères d’Europe et des pays limitrophes. Abeille 4: 1–131.
- Masumoto K. 1976a. A revision of the Coprophagid-beetles from Formosa (2). Elytra 4(1): 1–8.
- Masumoto K. 1976b. A revision of the Coprophagid-beetles from Formosa (3). Elytra 4(2): 25–30.
- Masumoto K, Tsai J-F, Ochi T. 2006. A revisional study of the Taiwanese Scarabaeinae (Coleoptera, Scarabaeidae). Part 3. Descriptions of a new species and a male of known species, lectotype designation, and downgrading of an Onthophagus species. Elytra 34(1): 139–144.
- Matsumura S. 1938. Onthophagid-insects from Formosa. Insecta Matsumurana 12(2–3): 53–63.
- Megerle von Mühlfeld JK. 1803. Appendix ad catalogum insectorum, quae mense Novembris MDCCCII Viennae Austriae auctionis lege vendita fuere. Privately published; Wien. 18 p
- Mittal IC. 1981. Distributional patterns in Scarabaeidae (Coleoptera): A Study on northwest India. Entomologische Blätter 77(1–2): 75–85.
- Mittal IC. 1986. Dung beetles attracted to human faeces. Entomologische Blätter 82(1–2): 55–64.
- Mittal IC. 1999. Annotated list of scarab fauna (Scarabaeidae: Coleoptera) of western Uttar Pradesh (U. P.) India. Annals of Entomology 17(2): 25–43.
- Mittal IC. 2000. Survey of scarabaeid (Coleoptera) fauna of Himachal Pradesh (India). Journal of Entomological Research 24(3): 259–269.
- Mittal IC, Jain R. 2015. A checklist Of Indian dung beetles (Coleoptera: Scarabaeidae). Indian Journal of Entomology 77(4): 383–404. https://doi.org/10.5958/0974-8172.2015.00076.0
- Miwa Y. 1930. An enumeration of the coprophagid-Coleoptera from Formosa with a table of the geographical distribution. Insecta Matsumurana 4(4): 163–180.
- Montreuil O. 2017. New records of Geotrupidae and Scarabeidae dung beetles (Insecta, Coleoptera) from Iran. Journal of Entomological Society of Iran 36(4): 259–277.
- Montrouzier X. 1855. Essai sur la Faune de l’Ile de Woodlark ou Moiou. Annales de la Société Entomologique de France 3(8): 229–308.
- Motschulsky V de. 1859. Sur les Collections Coléoptèrologiques de Linné et de Fabricius (continuation). XXII. Lamellicornes. Études Entomologiques 8: 147–168.
- Nakane T. 1956. On the Scarabaeidae of Japan (IV). The Entomological Review of Japan 12: 53–57.
- Nomura S. 1973. Notes on the Coprophagous Lamellicornia from Taiwan. The Entomological Review of Japan 25(1–2): 37–52.
- Olivier G-A. 1789. Entomologie, ou histoire naturelle des insectes, avec leurs caractères génériques et spécifiques, leur description, leur synonymie, et leur figure enluminée. Coléoptères. Tome premier (1–8). Baudouin; Paris. xx + 433 p. https://doi.org/10.5962/bhl.title.61905 https://doi.org/10.5962/bhl.title.61905
- Olivier G-A. 1790. Encyclopédie méthodique, ou par ordre de matières; par une société de gens de lettres, de savans et d’artistes; précédée d’un vocabulaire universel, servant de table pour tout l’ouvrage, ornée des portraits de Mm. Diderot & d’Alembert, premiers éditeurs de l’Encyclopédie. Histoire naturelle [Dictionnaire encyclopédique de l’histoire naturelle]. Insectes. Tome cinquième. Panckoucke; Paris. 793 p
- Palestrini C. 1983. Note preliminari su un gruppo indo-africano di Onthophagus. Atti del XIII Congresso Nazionale Italiano di Entomologia, Sestriere–Torino: 71–74.
- Paulian R. 1945. Faune de L’Empire Français. III. Coléoptères Scarabéides de L’Indochine. Première Partie. Librairie Larose; Paris. 227 p
- Paulian R. 1948. Expedition to South-West Arabia 1937–8. 12. Coleoptera, Scarabaeidae: Troginae, Geotrupinae, Dynamopinae, Hybosorinae, Coprinae, Aphodiinae. British Museum (Natural History) 1: 141–155.
- Paulian R. 1980. Insects of Saudia Arabia. Coleoptera: Scarabaeoidea (1re contribution). Fauna of Saudi Arabia 2: 141–154.
- Petrovitz R. 1961. Zoologische Ergebnisse der Österreichischen Karakorum-Expedition 1958. II. Teil. Coleoptera: Scarabaeidae. Sitzungsberichte der Österreichischen Akademie der Wissenschaften. Mathematisch-naturwissenschaftliche Klasse 170(3–4): 101–118.
- Petrovitz R. 1967. Österreichische entomologische Expeditionen nach Persien und Afghanistan. Beiträge zur Coleopterologie. Teil XI. Lamellicornia. Annalen des Naturhistorischen Museums in Wien 70: 479–490.
- Poggi R. 2010. Gli “Annali” pubblicati dal Museo Civico di Storia Naturale “Giacomo Doria” di Genova: storia del periodico ed indici generali dei primi cento volumi (1870–2009). Annali del Museo Civico di Storia Naturale “Giacomo Doria” 101: 1–529.
- Rajagopal C, Anu A, Vinod KV. 2023. A checklist of dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) associated with the Vypin-Kadamakkudy Barrier Island System, Kerala, India. Journal of Asia-Pacific Biodiversity 16: 324–331.
- Rajan PD. 2006. Insecta: Coleoptera: Scarabaeoidea: Scarabaeidae (Dung beetles). p. 91–135. In: Thirumalai G, Krishnan S (eds.). Fauna of Biligiri Rangaswamy Temple Wildlife Sanctuary (Karnataka), Conservation Area Series, 27. Zoological Survey of India; Kolkata. 264 p.
- Rani A, Sanjayan KP. 2013. Community composition of dung beetle and population dynamics of Tiniocellus spinipes in a scrub Jungle ecosystem. Asian Journal of Biological and Life Sciences 2(3): 241–246.
- Redtenbacher L. 1868. Reise der Österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den befehlen des Commodore B. von Wüllerstorf-Urbair. Zoologischer Theil. Zweiter Band. Coleopteren. Mit Fünf Tafeln. Karl Gerold’s Sohn; Wien. iv + 249 p. + 5 pls
- Reitter E. 1892. Bestimmungs-Tabelle der Lucaniden und coprophagen Lamellicornen. XXIV. Heft. (Sonderabdruck aus dem XXX. Bande der Verhandlungen des naturforscheden Vereins in Brünn). Verlag des Verfassers; Brno. 230 p
- Ritsema C. 1888. Lijst der entomologische geschriften van Mr. J. W. van Lansberge, gevolgd door eene opgave der daarin beschreven nieuwe geslachten, ondergeslachten en soorten. Tijdschrift voor Entomologie 31: 201–234.
- Rossini M, Vaz-de-Mello F, Mann DJ. 2014. Onthophagus cervicornis Kirby, 1825, new synonym under Onthophagus dama (Fabricius, 1798) (Coleoptera, Scarabaeidae, Scarabaeinae). Zookeys 419: 111–115. https://doi.org/10.3897/zookeys.419.7849
- Sathiandran N, Thomas SK, Flemming AT. 2015. An illustrated checklist of dung beetles (Coleoptera: Scarabaeinae) from the Periyar Tiger Reserve, Kerala, India. Journal of Threatened Taxa 7(15): 8250–8258. https://doi.org/10.11609/jott.2466.7.15.8250-8258
- Sathiandran N, Vineesh PJ, Thomas SK. 2021. Dung preference and trophic association of dung beetles (Coleoptera: Scarabaeidae) in the moist forests of the South-western Ghats of the Indian subcontinent. Journal of Asia-Pacific Entomology 24: 739–748. https://doi.org/10.1016/j.aspen.2021.06.008
- Schaller JG. 1783. XIII. Neue Insekten. Abhandlungen der Hallischen Naturforschenden Gesellschaft 1: 217–328.
- Schatzmayr A. 1946. Gli Scarabeidi coprofagi della Libia e dell’Egitto. Atti della Società Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano 85(1–2): 40–84.
- Scheuern J. 1995. Taxonomie, Sexualdimorphismus und Neubeschreibungen orientalischer Onthophagus-Arten unter besonderer Berücksichtigung des Subgenus Colobonthophagus Balthasar 1935 (Coleoptera, Scarabaeidae). Entomologica Basiliensia 18: 413–453.
- Scheuern J. 1996. Neue und verkannte Onthophagus-Arten aus der orientalischen Region (Coleoptera: Scarabaeidae). New and misinterpreted Onthophagus-Species from the Oriental Region (Coleoptera: Scarabaeidae). Stuttgarter Beiträge zur Naturkunde. Serie A (Biologie) 542: 1–20.
- Schmitt M, Hübner H, Gaedike R. 1998. Nomina auctorum—Auflösung von Abkürzungen taxonomischer Autoren-Namen. Nova Supplementa Entomologica 11: 1–192.
- Schönherr CJ. 1806. Synonymia Insectorum, eder: Versuch einer Synonymie Aller bisher bekannten Insecten; nach Fabricii Systema Eleutheratorum geordnet. Erster Band. Eleutherata oder Käfer. Erster Theil. Lethrus – Scolytes. Heinr. A. Nordström; Stockholm. xxii + 293 p. + 3 pls https://doi.org/10.5962/bhl.title.66107
- Schoolmeesters P. 2023. World Scarabaeidae Database. In: Bánki O, Roskov Y, Döring M, Ower G, Hernández Robles DR, Plata Corredor CA, Stjernegaard Jeppesen T, Örn A, Vandepitte L, Hobern D, Schalk P, DeWalt RE, Keping M, Miller J, Orrell T, Aalbu R, Abbott J, Adlard R, Adriaenssens EM, et al. (eds.). Catalogue of Life Checklist (Version 2023-05-30) https://doi.org/10.48580/dfsr-38g
- Sewak R. 1986a. Male genitalia of Indian Onthophagus Latr. and its taxonomic importance. Journal of Animal Morphology and Physiology 33(1–2): 135–140.
- Sewak R. 1986b. On a collection of dung beetles (Coleoptera: Scarabaeidae: Coprinae) from Rajasthan, India. Oikoassay 3(1): 11–16.
- Sewak R. 2003. Dung Beetles (Coleoptera: Scarabaeidae: Coprinae) of India with special reference to Arunachal Pradesh, Uttar Pradesh, Gujarat and Rajasthan. p. 249–287. In: Gupta RK (ed.). Advancements in insect biodiversity. Agrobios; Jodhpur. 337 p
- Sewak R. 2004. Insecta: Coleoptera: Scarabaeidae: Coprinae (dung beetles). p. 105–125. In: Alfred JRB (ed.). Fauna of Gujarat, State Fauna Series 8 (part 2). Zoological Survey of India; Kolkata. 427 p
- Sewak R. 2005. Dung beetles (Coleoptera: Scarabaeaidae: Coprinae) of the Thar Desert of Rajasthan. p. 143–148. In: Tyagi BK, Baqri QH (eds.). Changing faunal ecology in Thar Desert. Scientific Publishers; Jodhpur. 367 p
- Sewak R. 2006. Insecta: Coleoptera: Scarabaeidae: Coprinae (dung beetles). p. 191–224. In: Alfred JRB (ed.). Fauna of Arunachal Pradesh, State Fauna Series. 13 (part 2). Zoological Survey of India; Kolkata. 518 p
- Sewak R. 2009a. Dung beetles (Coleoptera:Scarabaeidae: Coprinae) of Thar Desert of Gujarat. Records of the Zoological Survey of India, Occasional Paper 295: 1–48.
- Sewak R. 2009b. Dung beetles (Coleoptera: Scarabaeidae: Coprinae) of Rajasthan. Records of the Zoological Survey of India, Occasional Paper 296: 1–106.
- Sewak R. 2010. Insecta: Coleoptera: Scarabaeidae. p. 93–118. In: Kumar S, Bohra P. Fauna of Ranthambhore Tiger Reserve, Rajasthan. Conservation Area Series 43. Zoological Survey of India; Kolkata. 229 p
- Sewak R, Yadva VK. 1991. Dung beetles (Coleoptera: Scarabaeidae: Coprinae) from five districts of Western Uttar Pradesh. Oikoassay 8(1): 25–27.
- Ship JW. 1895. Notes on Onthophagus, Latr.; with corrections of nomenclature, and a description of a new genus. The Entomologist 28: 178–179. https://doi.org/10.5962/bhl.part.11235
- Shrestha PK. 2005. Notes on Scarabaeinae fauna, their distribution and status in Nepal. Journal of Natural History Museum 22: 99–120.
- Siddiqui H, Ahmed Z, Khatri I. 2014. Distributional notes and new records for the dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) of Pakistan. Pakistan Journal of Zoology 46(2): 295–307.
- Siddiqui H, Kamaluddin S. 2011. The genus Onthophagus Latreille, 1802 (sensu lato) (Coleoptera: Scarabaeidae) in Pakistan. International Journal of Biology and Biotechnology 8(3): 347–352.
- Singh AP. 2020. Scarab beetles (Coleoptera: Scarabaeidae) of Dildar-Karnah (Kupwara, Jammu and Kashmir). Munis Entomology & Zoology 15(2): 521–525.
- Singh AP, Mahajan S, Uniyal V, Mondal R. 2017. First report on Scarabaeid (Coleoptera) fauna of Sahaspur, Uttarakhand. Journal of Entomology and Zoology Studies 5(6): 17–22.
- Sobhana KA, Thomas SK, Benny TM. 2013. Checklist of dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) associated with a dry deciduous forest in the South Western Ghats. Exapoda (Insecta indica) 20(1–2): 94–97.
- Streubel AV. 1846. Das Thierreich geordnet nach seiner Organisation, als Grundlage der Naturgeschichte der Thiere und als Einleitung in die vergleichende Anatomie. Vom Freiherrn Georg v. Cuvier. Nach der zweiten, vermehrten, Ausgabe frei ins Deutsche übersetzt und durch Zusätze sowohl dem heutigen Standpunkte der Wissenschaft angepasst als auch für den Selbstunterricht eingerichtet. Erster Theil. G. Reimer; Berlin. xvii + [1] + 972 p
- Sturm J. 1802. Abbildungen zu Karl Illiger’s Uebersetzung von Olivier’s Entomologie oder Naturgeschichte der Insecten mit inhren Gattungs- und Artmerkmalen ihrer Beschreibung und Synonymie. Käfer. Erster Theil. Mit vier und funfzig illuminirten Kupfertafeln; Nuremberg. iv + 136 p https://doi.org/10.5962/bhl.title.12457
- Sturm J. 1826. Catalog meiner Insecten-Sammlung. Erster Theil. Käfer. Mit 4 ausgemalten Kupfertafeln. Nürnberg. 207 p https://doi.org/10.5962/bhl.title.15090
- Sturm J. 1843. Catalog der Kaefer-Sammlung von Jacob Sturm. Mit 6 ausgemalten Kupfertafeln; Nuremberg. 386 p https:// doi.org/10.5962/bhl.title.37837
- Tarasov SI, Solodovnikov AY. 2011. Phylogenetic analyses reveal reliable morphological markers to classify megadiversity in Onthophagini dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae). Cladistics 27: 1–39. https://doi.org/10.1111/j.1096-0031.2011.00351.x
- Thakare V, Zade V, Chandra K, Gupta D. 2012. Scarab beetles (Coleoptera: Scarabaeoidea) of Melghat Tiger Reserve, Central India. Academic Journal of Entomology 5(2): 73–80.
- Thomas SK, Sathiandran N, Vinod KV. 2011. Faunal survey, endemism and possible species loss of Scarabaeinae (Coleoptera: Scarabaeidae) in the western slopes of the moist South Western Ghats, South India. Zootaxa 2830: 29–38.
- Thomas SK, Vinod KV, Vineesh PJ. 2006. Guild structure. diversity and succession of dung beetles associated with Indian elephant dung in South Western Ghats forests. Journal of Insect Science 6(17): 1–12. https://doi.org/10.1673/2006_06_17.1
- Thomas SK, Vinod KV, Vineesh PJ. 2007. Succession of dung beetles (Scarabaeinae: Coleoptera) in the dung pats of gaur, Bos gaurus H. Smith (Artiodactyla: Bovidae), from the moist deciduous forests of Southern Western Ghats. Biosystematica 1(1): 59–69. https://doi.org/10.1673/2006_06_17.1
- Van Harten A. 2005. The insects of the United Arab Emirates. A checklist of published Records. Dar Al Ummah Printing; Abu Dhabi. 87 p
- Venugopal KS, Thomas SK, Flemming AT. 2012. Diversity and community structure of dung beetles (Coleoptera: Scarabaeinae) associated with semi-urban fragmented agricultural land in the Malabar coast in southern India. Journal of Threatened Taxa 4(7): 2685–2692. https://doi.org/10.11609/JoTT.o3074.2685-92
- Vinod KV, Thomas SK. 2006. Species composition and community structure of dung beetles attracted to dung of gaur and elephant in the moist forests of South Western Ghats. Journal of Insect Science 7(56): 1–14. https://doi.org/10.1673/031.007.5601
- Walker FA. 1858. XXIII. Characters of some apparently undescribed Ceylon Insects. Annals and Magazine of Natural History 3(2): 202–209. https://doi.org/10.1080/00222935808697009
- Westermann BW. 1821. Schreiben des Herrn B. W. Westermann in Copenhagen, an den Profess. Wiedemann in Kiel. Aus dem Englischen überfetzt mit einigen Anmerkungen vom Empfänger. Magazin der Entomologie 4: 411–427.
- Westwood JO. 1842. Natural history of the insects of India, containing upwards of two hundred and twenty figures and descriptions, by E. Donovan, F. L. S. & W. S. A new edition, brought down to the present state of the science, with systematic characters of each species, synonyms, index, and other additional matter. Henry G. Bohn; London. 99 p
- Winkler A. 1929. Catalogus Coleopterorum regionis palaearcticae (1924–1932). Scarabaeidae. Wien 9: 1009–1136.
- Ziani S. 2015. Onthophagus (s. l.) spinifex (Fabricius, 1781). p. 19. In: Ziani S, Bezděk A, Branco T, Hillert O, Jákl S, Král D, Mantic M, Rößner E, Sehnal R. New country records of Scarabaeoidea (Coleoptera) from the Palaearctic Region. Insecta Mundi 0409: 1–36.
- Ziani S. 2020a. Some phonetic remarks on the pronunciation of the name Onthophagus (Insecta: Coleoptera: Scarabaeidae: Onthophagini). Quaderno di Studi e Notizie di Storia Naturale della Romagna 52: 219–221.
- Ziani S. 2020b. Preliminary considerations on the genus-group Furconthophagus and the Palaearctic species assigned to it in the literature (Coleoptera: Scarabaeidae: Onthophagini). Acta Societatis Zoologicae Bohemicae 84: 99–109.
- Aldhafe, HM, Barbero E. 2019. An overview of the Onthophagini from the Arabian Peninsula (Coleoptera: Scarabaeoidea: Scarabaeidae). Zootaxa 4658(1): 1–36. https://doi.org/10.11646/zootaxa.4658.1.1
- Ziani S, Bezděk A. 2016. Scarabaeidae, subfamily Scarabaeinae, tribe Onthophagini. p. 180–204. In: Löbl I, Löbl D. (eds.). Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea – Scirtoidea – Dascilloidea – Buprestoidea – Byrrhoidea. Revised and updated edition. Brill; Leiden, Boston. 983 p
- Ziani S, Gudenzi I. 2006. Studies on Palearctic Onthophagus associated with burrows of small mammals. I. O. furciceps, O. kindermanni, O. vitulus and closely related species. Bollettino della Società Entomologica Italiana 138(3): 207–248.
- Ziani S, Gudenzi I. 2009. Studies on palearctic Onthophagus associated with burrows of small mammals. III. O. aerarius and closely related species (Coleoptera Scarabaeidae). Bollettino della Società Entomologica Italiana 141(1): 29–44.
- Ziani S, Keith D. 2023. 3rd contribution to the knowledge of Arabian Onthophagini (Coleoptera: Scarabaeidae). Onthophagus aeneopiceus d’Orbigny, 1902, a new record for Yemen and for the Palaearctic ecozone. Insecta Mundi 0974: 1–4.
- Zimsen E. 1964. The type material of I. C. Fabricius. Munksgaard; Copenhagen. 656 p
- Zunino M. 1975. Revisione delle specie paleartiche del sottogenere Onthophagus (sensu strictu) Latr. (Coleoptera, Scarabaeoidea). I tipi di H. d’Orbigny, A. Raffray e A. Boucomont nel Muséum National d’Histoire Naturelle di Parigi. Bollettino del Museo di Zoologia dell’Università di Torino 7: 151–194.
- Zunino M. 1981. Insects of Saudi Arabia. Coleoptera: Fam. Scarabaeidae, Tribe Onthophagini. Fauna of Saudi Arabia 3: 408–416.
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