Abstract

Monogedania, a new fossil monotypic genus of predatory midge (Diptera: Ceratopogonidae) is described from Eocene Baltic amber and its position within the Monohelea complex is discussed. We dis¬covered that the membranous portion of the aedeagus is extended in Monogedania clunipes (Loew), new combination, which suggests that the aedeagus of some extinct predatory midges can be penis-like. The Eocene Monohelea baltica Szadziewski, is transferred to the genus Schizohelea Kieffer, new combination, and, the previously unknown female is described, and key characters are included in color photographs of its entire habitus, head, distal hind tarsomeres and claws.

References

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• Sinclair BJ, Borkent A, Wood DM. 2007. The male genital tract and aedeagal components of the Diptera with a discussion of their phylogenetic significance. Zoological Journal of the Linnean Society 150: 711–742. 
• Sontag E, Szadziewski R. 2011. Biting midges (Diptera: Ceratopogonidae) in Eocene Baltic amber from the Rovno region (Ukraine). Polish Journal of Entomology 80: 779–800. 
• Spinelli GR, Grogan WL Jr. 1998. A revision of the Neotropical predaceous midges of Brachypogon (Brachypogon) Kieffer (Diptera: Ceratopogonidae). Insecta Mundi 12 (1–2): 59–79. 
• Spinelli GR, Huerta H. 2015. Four new species of Mesoamerican biting midges of the genus Culicoides (Diptera: Ceratopogonidae). Acta Entomologica Musei Nationalis Pragae 55: 811–824. 
• Spinelli GR, Marino PI, Huerta H. 2015. Revision of the Neotropical species of the subgenus Atrichopogon (Psilokempia) (Diptera: Ceratopogonidae). Zootaxa 4003(1): 1–64. 
• Spinelli GR, Wirth WW. 1989. Las especies Neotropical del genero Bezzia (Diptera: Ceratopogonidae). II. Los grupos nobilis y punctipennis. Revista de la Sociedad Entomológica Argentina 45: 109–129. 
• Szadziewski R. 1988. Biting midges (Diptera, Ceratopogonidae) from Baltic amber. Polskie Pismo Entomologiczne 58: 3–283. 
• Szadziewski R. 1993. Biting midges (Diptera, Ceratopogonidae) from Miocene Saxonian amber. Acta Zoologica Cracoviensia 35: 603–656. 
• Szadziewski R. 1996. Biting midges from Lower Cretaceous amber of Lebanon and Upper Cretaceous Siberian amber of Taimyr (Diptera, Ceratopogonidae). Studia Dipterologica 3: 23–86. 
• Szadziewski R. 2018. Biting midges (Diptera: Ceratopogonidae) as indicators of biostratigraphy, ecological reconstructions and identification of amber deposits. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 107: 219–230. 
• Szadziewski R, Grogan WL Jr. 1998. Biting midges from Dominican amber. IV. Species of the tribes Dasyheleini and Forcipomyiini (Diptera: Ceratopogonidae). Polskie Pismo Entomologiczne (Polish Journal of Entomology) 67: 255–290. 
• Szadziewski R, Dominiak P, Withers P. 2020. Two new species of biting midges from France and Algeria (Diptera: Ceratopogonidae). Annales Zoologici 70: 113–120. 
• Wirth WW. 1953. American biting midges of the heleid genus Monohelea. Proceedings of the United States National Museum 103: 135–154. 
• Wirth WW, Blanton FS. 1972. A new Patagonian biting midge of the genus Monohelea, Isthmohelea (Diptera: Ceratopogonidae). Florida Entomologist 55: 173–176. 
• Wirth WW, Grogan WL Jr. 1988. The predaceous midges of the world (Diptera: Ceratopogonidae; Tribe Ceratopogonini). Flora and fauna handbook 4. E. J. Brill; New York. 160 p