A new genus of predatory midge in the Monohelea complex from Eocene Baltic amber (Diptera: Ceratopogonidae)
Ryszard Szadziewski
Department of Invertebrate Zoology and Parasitology University of Gdańsk, Wita Stwosza 59, 80-308 Gdańsk, Poland
William L., Jr. Grogan
Florida State Collection of Arthropods Florida Dept. of Agriculture & Consumer Services, Gainesville, FL 32614-7100 U.S.A.
Elżbieta Sontag
Department of Invertebrate Zoology and Parasitology University of Gdańsk, Wita Stwosza 59, 80-308 Gdańsk, Poland
Błażej Bojarski
Department of Invertebrate Zoology and Parasitology University of Gdańsk, Wita Stwosza 59, 80-308 Gdańsk, Poland
- Keywords
- Ceratopogoninae, Ceratopogonini, aedeagus
Abstract
Monogedania, a new fossil monotypic genus of predatory midge (Diptera: Ceratopogonidae) is described from Eocene Baltic amber and its position within the Monohelea complex is discussed. We dis¬covered that the membranous portion of the aedeagus is extended in Monogedania clunipes (Loew), new combination, which suggests that the aedeagus of some extinct predatory midges can be penis-like. The Eocene Monohelea baltica Szadziewski, is transferred to the genus Schizohelea Kieffer, new combination, and, the previously unknown female is described, and key characters are included in color photographs of its entire habitus, head, distal hind tarsomeres and claws.
References (43)
- Alwin-Kownacka A, Szadziewski R, Szwedo, J. 2016. Biting midges of the tribe Ceratopogonini (Diptera: Ceratopogonidae) from the Middle East, with keys and descriptions of new species. Zootaxa 4079: 551–572.
- Borkent A. 1995. Biting midges in the Cretaceous amber of North America (Diptera: Ceratopogonidae). Backhuis Publishers; Leiden. 237 p
- Borkent A, Dominiak P. 2020. Catalog of biting midges of the World (Diptera: Ceratopogonidae). Zootaxa 4787: 1–377.
- Ingram A, Macfie JWS. 1931. Ceratopogonidae. p. 155–232. In: Diptera of Patagonia and South Chile. Part II. Fasc. 4. British Museum (Nat. Hist.); London, United Kingdom. 331 p
- Kieffer JJ. 1917. Chironomides d’Amerique conservés au Musée National Hongrois de Budapest. Annales Historico-Naturales Musei Nationalis Hungarici 15: 292–364.
- Loew H. 1850. Über den Bernstein und die Bernsteinfauna. Program der Königlichen Reaschule zu Meseritz 1: 3–44.
- Meigen JW. 1804. Klassifikazion und Beschreibung der europäischen zweiflügeligen Insekten (Diptera Linn.). Braunschweig 1: 1–152.
- Meunier F. 1904. Monographie des Cecidomyidae, des Sciaridae, des Mycetophilidae et des Chironomidae de l’ambre de la Baltique. Annales de la Société Scientifique de Bruxelles 28: 12–264.
- Peñalver E, Arillo A, Szadziewski R, Jeffrey DS. 2021. Biting midges (Diptera: Ceratopogonidae) from the middle Eocene Anglesea amber (Australia) originated in a subpolar greenhouse earth. Historical Biology 2021: 1–11. https://doi.org/10.1080/08912963.2021.1924700
- Ronderos MM, Spinelli GR, Grogan WL Jr. 2017. The Neotropical species of the predaceous midge genus Austrohelea Wirth & Grogan (Diptera: Ceratopogonidae). Zootaxa 4276: 255–269.
- Santarém MC, Borkent A, Felippe-Bauer ML. 2019. Taxonomic revision of Neotropical Downeshelea Wirth and Grogan predaceous midges (Diptera: Ceratopogonidae). Insects 11(1), 9: 1–94.
- Schmidt AR, Kaulfuss UJ, Bannister M, Baranov V, Beimforde C, Bleile N, Borkent A, Busch A, Conran JG, Harvey MS, Kennedy EM, Kerr PH, Kettunen E, Kiecksee AP, Lengeling F, Lindqvist JK, Maraun M, Mildenhall D, Perrichot V, Rikkinen J, Sadowski E-M, Seyfullah LJ, Stebner F, Szwedo J, Ulbrich P, Lee DE. 2018. Amber inclusions from New Zealand. Gondwana Research 56: 135–146.
- Sinclair BJ, Borkent A, Wood DM. 2007. The male genital tract and aedeagal components of the Diptera with a discussion of their phylogenetic significance. Zoological Journal of the Linnean Society 150: 711–742.
- Sontag E, Szadziewski R. 2011. Biting midges (Diptera: Ceratopogonidae) in Eocene Baltic amber from the Rovno region (Ukraine). Polish Journal of Entomology 80: 779–800.
- Spinelli GR, Grogan WL Jr. 1998. A revision of the Neotropical predaceous midges of Brachypogon (Brachypogon) Kieffer (Diptera: Ceratopogonidae). Insecta Mundi 12 (1–2): 59–79.